中国农业科技导报 ›› 2024, Vol. 26 ›› Issue (5): 52-64.DOI: 10.13304/j.nykjdb.2022.0869
卢登洋1(), 童盼盼1, 闫敏1, 鲍荆凯1, 刘鸣哲1, 夏怡蕾1, 吴翠云1,2(
)
收稿日期:
2022-10-13
接受日期:
2023-04-12
出版日期:
2024-05-15
发布日期:
2024-05-14
通讯作者:
吴翠云
作者简介:
卢登洋E-mail:1729854434@qq.com;
基金资助:
Dengyang LU1(), Panpan TONG1, Min YAN1, Jingkai BAO1, Mingzhe LIU1, Yilei XIA1, Cuiyun WU1,2(
)
Received:
2022-10-13
Accepted:
2023-04-12
Online:
2024-05-15
Published:
2024-05-14
Contact:
Cuiyun WU
摘要:
为鉴定和评价库尔勒香梨大果芽变的性状,为生产栽培和遗传育种提供理论依据,在2020—2021年对库尔勒香梨(对照)及其大果芽变的植物学形态、生物学特性、品质特性进行比较分析。结果表明,大果芽变果实呈倒卵形,库尔勒香梨呈纺锤形;大果芽变的叶片保卫细胞宽度和果肉细胞长度均显著增大;在果实动态发育过程中,大果芽变的单果重及果实含水率均显著高于库尔勒香梨,在果实成熟期,与库尔勒香梨相比,大果芽变果实的可溶性糖含量及糖酸比分别显著提高78.39%和84.54%,石细胞含量显著降低33.33%。经表型性状和品质特性分析,判定大果芽变为库尔勒香梨的优良芽变,可作为品种改良及新品种选育的亲本材料。
中图分类号:
卢登洋, 童盼盼, 闫敏, 鲍荆凯, 刘鸣哲, 夏怡蕾, 吴翠云. 库尔勒香梨大果芽变的鉴定与评价[J]. 中国农业科技导报, 2024, 26(5): 52-64.
Dengyang LU, Panpan TONG, Min YAN, Jingkai BAO, Mingzhe LIU, Yilei XIA, Cuiyun WU. Identification and Evaluation of Korla Pear Bud Sport with Larger Fruit Size[J]. Journal of Agricultural Science and Technology, 2024, 26(5): 52-64.
品种(系) Cultivar (strain) | 花芽萌芽期 Date of bud sprouting | 展叶期 Date of leaf flatting | 现蕾期 Date of budding stage | 初花期 Date of first bloom | 盛花期 Date of full bloom | 末花期 Date of flower dropping | 成熟期 Date of mature |
---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 3/29 | 4/13 | 4/9 | 4/10 | 4/10 | 4/18 | 9/21 |
芽变 Bud sport | 3/31 | 4/14 | 4/9 | 4/11 | 4/12 | 4/20 | 9/20 |
表1 库尔勒香梨及其芽变的物候期 (m/d)
Table 1 Phenological stages of Korla pear and bud sport
品种(系) Cultivar (strain) | 花芽萌芽期 Date of bud sprouting | 展叶期 Date of leaf flatting | 现蕾期 Date of budding stage | 初花期 Date of first bloom | 盛花期 Date of full bloom | 末花期 Date of flower dropping | 成熟期 Date of mature |
---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 3/29 | 4/13 | 4/9 | 4/10 | 4/10 | 4/18 | 9/21 |
芽变 Bud sport | 3/31 | 4/14 | 4/9 | 4/11 | 4/12 | 4/20 | 9/20 |
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
树姿 Tree posture | 开张 Opened | 开张 Opened |
主干树皮特征 Main stem bark features | 片状剥落 Flaky spalling | 光滑 Smooth |
1年生枝皮孔数量 Number of lenticels on annual branches | 少 Little | 多 More |
叶芽姿态 Leaf bud posture | 贴生 Poster | 贴生 Poster |
芽托大小 Bud holder size | 中 Medium | 小 Small |
连续结果能力 Continuous results capability | 强 Stronger | 强 Stronger |
树势 Tree strength | 强 Stronger | 强 Stronger |
成枝能力 Branching capability | 强 Stronger | 强 Stronger |
叶芽顶端特征 Apical features of leaf buds | 尖 Pointed | 尖 Pointed |
花芽茸毛 Flower buds velvet | 有 Available | 有 Available |
嫩枝茸毛 Tender fuzzy shoots | 无 None | 无 None |
表2 库尔勒香梨及其芽变树性特征
Table 2 Characteristics of Korla pear and bud sport
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
树姿 Tree posture | 开张 Opened | 开张 Opened |
主干树皮特征 Main stem bark features | 片状剥落 Flaky spalling | 光滑 Smooth |
1年生枝皮孔数量 Number of lenticels on annual branches | 少 Little | 多 More |
叶芽姿态 Leaf bud posture | 贴生 Poster | 贴生 Poster |
芽托大小 Bud holder size | 中 Medium | 小 Small |
连续结果能力 Continuous results capability | 强 Stronger | 强 Stronger |
树势 Tree strength | 强 Stronger | 强 Stronger |
成枝能力 Branching capability | 强 Stronger | 强 Stronger |
叶芽顶端特征 Apical features of leaf buds | 尖 Pointed | 尖 Pointed |
花芽茸毛 Flower buds velvet | 有 Available | 有 Available |
嫩枝茸毛 Tender fuzzy shoots | 无 None | 无 None |
品种(系)Cultivar (strain) | 叶片形状 Leaf shape | 叶基形状 Leaf base shape | 叶缘 Leaf margin | 叶背绒毛 Dorsal leaf tomentum | 叶尖形状 Leaf tip shape | 芒刺 Mansard | 托叶 Stipule | 叶姿 Leaf stance | 叶面状态 Foliar condition |
---|---|---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 圆形 Circular | 圆形 Circular | 锐锯齿 Sharp serrations | 无 None | 渐尖 Acuminate | 有 Available | 无 None | 水平 Horizontal | 波浪 Wave |
芽变 Bud sport | 卵圆形 Ovoid | 宽楔形 Broad wedge | 复锯齿 Compound serrations | 无 None | 渐尖 Acuminate | 有 Available | 无 None | 斜向上 Slanted upward | 波浪 Wave |
表3 库尔勒香梨及其芽变的叶片质量性状
Table 3 Leaf quality traits of Korla pear and bud sport
品种(系)Cultivar (strain) | 叶片形状 Leaf shape | 叶基形状 Leaf base shape | 叶缘 Leaf margin | 叶背绒毛 Dorsal leaf tomentum | 叶尖形状 Leaf tip shape | 芒刺 Mansard | 托叶 Stipule | 叶姿 Leaf stance | 叶面状态 Foliar condition |
---|---|---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 圆形 Circular | 圆形 Circular | 锐锯齿 Sharp serrations | 无 None | 渐尖 Acuminate | 有 Available | 无 None | 水平 Horizontal | 波浪 Wave |
芽变 Bud sport | 卵圆形 Ovoid | 宽楔形 Broad wedge | 复锯齿 Compound serrations | 无 None | 渐尖 Acuminate | 有 Available | 无 None | 斜向上 Slanted upward | 波浪 Wave |
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
叶长 Leaf length/mm | 55.87±7.53 b | 63.67±7.37 a |
叶宽 Leaf width/mm | 38.79±4.06 a | 41.42±3.30 a |
叶形指数 Leaf shape index | 1.46±0.26 a | 1.55±0.22 a |
叶面积 Leaf area/mm2 | 1 486.47±232.68 b | 1 858.71±395.33 a |
叶柄长 Petiole length/mm | 26.45±6.57 a | 28.04±7.45 a |
叶柄宽 Petiole width/mm | 3.51±1.32 a | 4.77±6.63 a |
叶柄长/宽 Petiole length/width | 8.27±2.76 a | 7.70±2.30 a |
叶绿素参考值 Chlorophyll reference values | 5.77±1.39 a | 5.58±0.22 a |
表4 库尔勒香梨及其芽变的叶片数量性状
Table 4 Leaf traits between Korla pear and bud sport
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
叶长 Leaf length/mm | 55.87±7.53 b | 63.67±7.37 a |
叶宽 Leaf width/mm | 38.79±4.06 a | 41.42±3.30 a |
叶形指数 Leaf shape index | 1.46±0.26 a | 1.55±0.22 a |
叶面积 Leaf area/mm2 | 1 486.47±232.68 b | 1 858.71±395.33 a |
叶柄长 Petiole length/mm | 26.45±6.57 a | 28.04±7.45 a |
叶柄宽 Petiole width/mm | 3.51±1.32 a | 4.77±6.63 a |
叶柄长/宽 Petiole length/width | 8.27±2.76 a | 7.70±2.30 a |
叶绿素参考值 Chlorophyll reference values | 5.77±1.39 a | 5.58±0.22 a |
品种(系) Cultivar (strain) | 上表皮厚度 Upper skin thickness/μm | 下表皮厚度 Lower skin thickness/μm | 叶片厚度 Leaf thickness/μm | 栅栏组织 厚度 Fence tissue thickness/μm | 海绵组织 厚度 Spongy tissue thickness/μm | 栅海比 Fence to spongy ratio | 组织结构疏松度 Looseness of tissue structure | 组织结构紧密度Tightness of tissue structure |
---|---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 20.04±6.07 a | 12.85±3.46 a | 277.98±11.81 a | 130.75±22.52 a | 104.31±23.23 a | 1.31±0.37 a | 0.37±0.08 a | 0.47±0.08 a |
芽变 Bud sport | 17.84±4.77 a | 13.09±2.33 a | 282.78±9.77 a | 131.76±19.47 a | 111.54±16.74 a | 1.21±0.27 a | 0.39±0.06 a | 0.47±0.07 a |
表5 库尔勒香梨及其芽变的叶片解剖结构
Table 5 Anatomical structure of the leaves of Korla pear and bud sport
品种(系) Cultivar (strain) | 上表皮厚度 Upper skin thickness/μm | 下表皮厚度 Lower skin thickness/μm | 叶片厚度 Leaf thickness/μm | 栅栏组织 厚度 Fence tissue thickness/μm | 海绵组织 厚度 Spongy tissue thickness/μm | 栅海比 Fence to spongy ratio | 组织结构疏松度 Looseness of tissue structure | 组织结构紧密度Tightness of tissue structure |
---|---|---|---|---|---|---|---|---|
库尔勒香梨 Korla pear | 20.04±6.07 a | 12.85±3.46 a | 277.98±11.81 a | 130.75±22.52 a | 104.31±23.23 a | 1.31±0.37 a | 0.37±0.08 a | 0.47±0.08 a |
芽变 Bud sport | 17.84±4.77 a | 13.09±2.33 a | 282.78±9.77 a | 131.76±19.47 a | 111.54±16.74 a | 1.21±0.27 a | 0.39±0.06 a | 0.47±0.07 a |
品种(系) Cultivar (strain) | 气孔数目 Number of stomata | 气孔密度 Stomatal density/mm-2 | 保卫细胞长度 Defending cell length/μm | 保卫细胞宽度 Defending cell width/μm | |
---|---|---|---|---|---|
范围 Range | 平均值 Mean | ||||
库尔勒香梨 Korla pear | 6~14 | 9.97±1.92 a | 112.56±21.72 a | 145.05±16.56 a | 105.09±17.28 b |
芽变 Bud sport | 6~13 | 9.60±1.84 a | 108.42±20.74 a | 139.37±16.72 a | 121.73±12.88 a |
表6 库尔勒香梨及其芽变的叶片气孔性状
Table 6 Stomata in leaves of Korla pear and bud sport
品种(系) Cultivar (strain) | 气孔数目 Number of stomata | 气孔密度 Stomatal density/mm-2 | 保卫细胞长度 Defending cell length/μm | 保卫细胞宽度 Defending cell width/μm | |
---|---|---|---|---|---|
范围 Range | 平均值 Mean | ||||
库尔勒香梨 Korla pear | 6~14 | 9.97±1.92 a | 112.56±21.72 a | 145.05±16.56 a | 105.09±17.28 b |
芽变 Bud sport | 6~13 | 9.60±1.84 a | 108.42±20.74 a | 139.37±16.72 a | 121.73±12.88 a |
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
花瓣形状 Petal shapes | 圆形 Circular | 卵圆形 Ovoid |
花瓣位置 Petal position | 邻接 Adjacent | 邻接 Adjacent |
花蕾颜色 Flower bud colour | 浅粉色 Pale pink | 浅粉色 Pale pink |
柱头位置 Column head position | 高于花药 Higher than anthers | 与花药等高 As high as anthers |
花瓣基部茸毛 Petal base velutinous | 有 Available | 无 None |
花药颜色 Anther colour | 粉红色 Pink colour | 粉红色 Pink colour |
花序花朵数 Number of flowers per inflorescence | 8.23±1.98 a | 6.07±1.11 b |
雌蕊数 Number of pistils | 5.00±0.00 a | 5.00±0.00 a |
雄蕊数 Number of stamens | 19.87±1.33 a | 20.53±1.11 a |
花冠直径 Corolla diameter/mm | 23.55±3.30 a | 26.15±2.96 a |
花瓣横径 Petal diameter/mm | 8.23±1.98 a | 6.07±1.11 b |
花瓣纵经 Petal longitudinal diameter/mm | 10.56±1.51 a | 11.12±1.53 a |
单药花粉量 Pollen number per anther | 7 228.57±450.21 a | 3 535.71±255.75 b |
花粉生活力Pollen viability/% | 79.26±0.05 b | 86.68±0.05 a |
表7 库尔勒香梨及其芽变的花性状
Table 7 Flowering organ morphology between Korla pear and bud sport
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
花瓣形状 Petal shapes | 圆形 Circular | 卵圆形 Ovoid |
花瓣位置 Petal position | 邻接 Adjacent | 邻接 Adjacent |
花蕾颜色 Flower bud colour | 浅粉色 Pale pink | 浅粉色 Pale pink |
柱头位置 Column head position | 高于花药 Higher than anthers | 与花药等高 As high as anthers |
花瓣基部茸毛 Petal base velutinous | 有 Available | 无 None |
花药颜色 Anther colour | 粉红色 Pink colour | 粉红色 Pink colour |
花序花朵数 Number of flowers per inflorescence | 8.23±1.98 a | 6.07±1.11 b |
雌蕊数 Number of pistils | 5.00±0.00 a | 5.00±0.00 a |
雄蕊数 Number of stamens | 19.87±1.33 a | 20.53±1.11 a |
花冠直径 Corolla diameter/mm | 23.55±3.30 a | 26.15±2.96 a |
花瓣横径 Petal diameter/mm | 8.23±1.98 a | 6.07±1.11 b |
花瓣纵经 Petal longitudinal diameter/mm | 10.56±1.51 a | 11.12±1.53 a |
单药花粉量 Pollen number per anther | 7 228.57±450.21 a | 3 535.71±255.75 b |
花粉生活力Pollen viability/% | 79.26±0.05 b | 86.68±0.05 a |
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
果实形状 Fruit shape | 纺锤形 Spindle-shaped | 倒卵形 Obovate-shaped |
果皮底色 Peel base colour | 绿黄色 Green and yellow | 绿黄色 Green and yellow |
果面光滑度 Smoothness of fruit surface | 平滑 Smooth | 平滑 Smooth |
果面盖色 Fruit cover color | 橘红色 Red-orange | 橘红色 Red-orange |
着色程度 Degree of colouring | 部分着色 Partially coloured | 部分着色 Partially coloured |
着色类型 Type of colouring | 条红 Striped red | 条红 Striped red |
果锈数量Number of fruit rust | 无或极少 None or very few | 无或极少 None or very few |
果锈位置 Fruit rust location | 萼端 Calyx ends | 梗端 Stem ends |
棱沟 Furrows | 有 Available | 有 Available |
蜡质 Waxy | 无或极少 None or very few | 少 Little |
果梗基部 Base of fruit stem | 有 Available | 有 Available |
果梗姿态 Fruit stem posture | 斜生 Oblique growth | 斜生 Oblique growth |
梗洼深度 Depth of peduncle | 中 Medium | 浅 Light |
梗洼广度 Breadth of peduncle | 广 Broad | 广 Broad |
萼洼深度 Calyx pits depth | 浅 Light | 深 Depth |
萼洼广度 Calyx pits breadth | 广 Broad | 广 Broad |
萼洼状态 Calyx pits state | 肋状 Costal | 平滑 Smooth |
萼片状态 Calyx condition | 宿存 Cumulus | 宿存 Cumulus |
萼片姿态 Calyx posture | 聚合 Polymerisation | 聚合 Polymerisation |
种子颜色 Seed colour | 褐色 Brown | 深褐色 Dark brown |
种子形状 Seed shape | 椭圆 Oval-shaped | 卵圆 Ovoid |
种子个数 Number of seeds | 9 | 9 |
表8 库尔勒香梨及其芽变的果实描述型性状
Table 8 Fruit traits between Korla pear and bud sport
性状 Trait | 库尔勒香梨 Korla pear | 芽变 Bud sport |
---|---|---|
果实形状 Fruit shape | 纺锤形 Spindle-shaped | 倒卵形 Obovate-shaped |
果皮底色 Peel base colour | 绿黄色 Green and yellow | 绿黄色 Green and yellow |
果面光滑度 Smoothness of fruit surface | 平滑 Smooth | 平滑 Smooth |
果面盖色 Fruit cover color | 橘红色 Red-orange | 橘红色 Red-orange |
着色程度 Degree of colouring | 部分着色 Partially coloured | 部分着色 Partially coloured |
着色类型 Type of colouring | 条红 Striped red | 条红 Striped red |
果锈数量Number of fruit rust | 无或极少 None or very few | 无或极少 None or very few |
果锈位置 Fruit rust location | 萼端 Calyx ends | 梗端 Stem ends |
棱沟 Furrows | 有 Available | 有 Available |
蜡质 Waxy | 无或极少 None or very few | 少 Little |
果梗基部 Base of fruit stem | 有 Available | 有 Available |
果梗姿态 Fruit stem posture | 斜生 Oblique growth | 斜生 Oblique growth |
梗洼深度 Depth of peduncle | 中 Medium | 浅 Light |
梗洼广度 Breadth of peduncle | 广 Broad | 广 Broad |
萼洼深度 Calyx pits depth | 浅 Light | 深 Depth |
萼洼广度 Calyx pits breadth | 广 Broad | 广 Broad |
萼洼状态 Calyx pits state | 肋状 Costal | 平滑 Smooth |
萼片状态 Calyx condition | 宿存 Cumulus | 宿存 Cumulus |
萼片姿态 Calyx posture | 聚合 Polymerisation | 聚合 Polymerisation |
种子颜色 Seed colour | 褐色 Brown | 深褐色 Dark brown |
种子形状 Seed shape | 椭圆 Oval-shaped | 卵圆 Ovoid |
种子个数 Number of seeds | 9 | 9 |
图1 库尔勒香梨和大果芽变的单果重、果实纵径、果实横径和果形指数注:同一时期中不同小写字母表示在P<0.05水平差异显著。
Fig. 1 Fruit weight, fruit longitudinal diameter, fruit transverse diameterand fruit shape index of Korla pear and bud sportNote: Different lowercase letters in same period indicate significant differences at P<0.05 level.
采样日期 Sampling date (m/d) | 品种(系) Cultivar(strain) | 亮度指标L* | 红绿指标a* | 黄蓝指标b* | 色饱和度C | 色泽比h | 色差△E |
---|---|---|---|---|---|---|---|
6/13 | 库尔勒香梨 Korla pear | 35.29±13.59 a | -10.89±2.20 a | 27.78±4.57 a | 15.33±5.05 a | -0.53±0.04 b | 66.84±9.60 a |
芽变 Bud sport | 26.84±5.16 b | -12.13±3.50 b | 28.82±4.87 a | 15.87±5.03 a | -0.57±0.11 a | 66.99±8.87 a | |
6/28 | 库尔勒香梨 Korla pear | 24.18±8.88 b | -8.66±2.05 a | 25.46±4.70 b | 11.50±3.46 b | -0.49±0.06 b | 73.36±10.76 a |
芽变 Bud sport | 35.43±6.56 a | -14.97±8.10 b | 42.67±18.11 a | 31.10±14.37 a | -0.55±0.45 a | 43.38±13.48 b | |
7/13 | 库尔勒香梨 Korla pear | 24.23±6.21 b | -9.92±2.24 a | 24.23±6.21 b | 30.44±4.16 b | 2.61±0.22 a | 74.90±4.34 a |
芽变 Bud sport | 35.52±13.21 a | -12.18±4.90 b | 26.84±5.16 a | 36.58±5.65 a | 2.58±0.63 b | 73.40±3.25 a | |
7/28 | 库尔勒香梨 Korla pear | 31.94±7.11 b | -10.94±3.82 a | 36.56±6.61 b | 16.41±2.86 b | -0.46±0.17 a | 68.78±9.53 a |
芽变 Bud sport | 55.14±5.73 a | -14.51±4.00 b | 45.08±7.88 a | 21.11±5.46 a | -0.44±0.62 b | 68.33±4.47 a | |
8/12 | 库尔勒香梨 Korla pear | 44.74±6.95 b | -11.60±4.31 a | 55.82±9.29 b | 24.33±3.74 b | -0.37±0.12 a | 62.23±4.53 a |
芽变 Bud sport | 58.09±7.93 a | -14.77±5.52 b | 76.97±6.03 a | 33.25±4.69 a | -0.33±0.27 b | 57.14±8.24 b | |
8/27 | 库尔勒香梨 Korla pear | 55.38±6.59 b | -10.08±10.43 a | 73.27±9.61 b | 31.30±4.64 b | -0.11±0.25 b | 56.86±2.92 a |
芽变 Bud sport | 77.46±14.19 a | -10.26±6.17 a | 76.46±8.36 a | 35.33±7.79 a | -0.20±0.32 a | 56.33±3.29 a | |
9/11 | 库尔勒香梨 Korla pear | 81.99±16.74 a | -12.76±7.51 b | 96.81±11.84 a | 52.93±14.80 a | -0.31±0.45 a | 71.34±13.92 a |
芽变 Bud sport | 82.15±8.65 a | 0.61±11.29 a | 102.17±11.96 a | 50.80±5.85 a | 0.03±0.24 a | 69.63±11.05 a | |
9/21 | 库尔勒香梨 Korla pear | 88.59±10.42 a | -10.08±10.43 b | 99.16±9.13 a | 77.48±13.27 a | -0.83±4.12 a | 85.42±199.64 a |
芽变 Bud sport | 88.96±10.47 a | 7.57±15.48 a | 105.99±10.76 a | 78.58±12.68 a | 0.17±0.32 a | 55.39±6.24 b |
表9 库尔勒香梨和芽变发育过程中表皮色泽的比较
Table 9 Comparison of epidermal colour during bud development in Korla pear and bud sport
采样日期 Sampling date (m/d) | 品种(系) Cultivar(strain) | 亮度指标L* | 红绿指标a* | 黄蓝指标b* | 色饱和度C | 色泽比h | 色差△E |
---|---|---|---|---|---|---|---|
6/13 | 库尔勒香梨 Korla pear | 35.29±13.59 a | -10.89±2.20 a | 27.78±4.57 a | 15.33±5.05 a | -0.53±0.04 b | 66.84±9.60 a |
芽变 Bud sport | 26.84±5.16 b | -12.13±3.50 b | 28.82±4.87 a | 15.87±5.03 a | -0.57±0.11 a | 66.99±8.87 a | |
6/28 | 库尔勒香梨 Korla pear | 24.18±8.88 b | -8.66±2.05 a | 25.46±4.70 b | 11.50±3.46 b | -0.49±0.06 b | 73.36±10.76 a |
芽变 Bud sport | 35.43±6.56 a | -14.97±8.10 b | 42.67±18.11 a | 31.10±14.37 a | -0.55±0.45 a | 43.38±13.48 b | |
7/13 | 库尔勒香梨 Korla pear | 24.23±6.21 b | -9.92±2.24 a | 24.23±6.21 b | 30.44±4.16 b | 2.61±0.22 a | 74.90±4.34 a |
芽变 Bud sport | 35.52±13.21 a | -12.18±4.90 b | 26.84±5.16 a | 36.58±5.65 a | 2.58±0.63 b | 73.40±3.25 a | |
7/28 | 库尔勒香梨 Korla pear | 31.94±7.11 b | -10.94±3.82 a | 36.56±6.61 b | 16.41±2.86 b | -0.46±0.17 a | 68.78±9.53 a |
芽变 Bud sport | 55.14±5.73 a | -14.51±4.00 b | 45.08±7.88 a | 21.11±5.46 a | -0.44±0.62 b | 68.33±4.47 a | |
8/12 | 库尔勒香梨 Korla pear | 44.74±6.95 b | -11.60±4.31 a | 55.82±9.29 b | 24.33±3.74 b | -0.37±0.12 a | 62.23±4.53 a |
芽变 Bud sport | 58.09±7.93 a | -14.77±5.52 b | 76.97±6.03 a | 33.25±4.69 a | -0.33±0.27 b | 57.14±8.24 b | |
8/27 | 库尔勒香梨 Korla pear | 55.38±6.59 b | -10.08±10.43 a | 73.27±9.61 b | 31.30±4.64 b | -0.11±0.25 b | 56.86±2.92 a |
芽变 Bud sport | 77.46±14.19 a | -10.26±6.17 a | 76.46±8.36 a | 35.33±7.79 a | -0.20±0.32 a | 56.33±3.29 a | |
9/11 | 库尔勒香梨 Korla pear | 81.99±16.74 a | -12.76±7.51 b | 96.81±11.84 a | 52.93±14.80 a | -0.31±0.45 a | 71.34±13.92 a |
芽变 Bud sport | 82.15±8.65 a | 0.61±11.29 a | 102.17±11.96 a | 50.80±5.85 a | 0.03±0.24 a | 69.63±11.05 a | |
9/21 | 库尔勒香梨 Korla pear | 88.59±10.42 a | -10.08±10.43 b | 99.16±9.13 a | 77.48±13.27 a | -0.83±4.12 a | 85.42±199.64 a |
芽变 Bud sport | 88.96±10.47 a | 7.57±15.48 a | 105.99±10.76 a | 78.58±12.68 a | 0.17±0.32 a | 55.39±6.24 b |
图3 库尔勒香梨及其芽变果实的可溶性糖、可滴定酸、可溶性固形物含量和糖酸比注:同一时期中不同小写字母表示在P<0.05水平差异显著。
Fig. 3 Content of soluble sugars, titratable acids, soluble solids and sugar acid ratio in Korla pear and bud sportNote: Different lowercase letters in same period indicate significant differences at P<0.05 level.
图4 库尔勒香梨及其芽变果实Vc、蛋白质含量、石细胞含量、含水率、总黄酮和总酚含量注:同一时期中不同小写字母表示在P<0.05水平差异显著。
Fig. 4 Contents of Vc, protein, stone cell, water, total flavonoids and total phenolic of Korla pear and bud sportNote:Different lowercase letters in same period indicate significant differences at P<0.05 level.
采样日期 Sampling date(m/d) | 细胞长度 Cell size/μm | 细胞数量 Cell number/μm-2 | ||
---|---|---|---|---|
库尔勒香梨 Korla pear | 芽变 Bud sport | 库尔勒香梨 Korla pear | 芽变 Bud sport | |
6/13 | 183.22±32.53 b | 230.02±27.15 a | 20.07±3.56 a | 19.10±2.25 a |
6/28 | 257.08±30.09 b | 396.01±71.51 a | 25.22±2.95 a | 24.69±4.46 a |
7/13 | 426.63±35.74 b | 490.32±84.17 a | 31.07±2.60 a | 24.47±4.20 b |
7/28 | 512.87±67.87 b | 569.32±63.54 a | 21.37±2.83 b | 24.08±2.69 a |
8/12 | 579.04±81.59 b | 660.16±109.95 a | 26.15±3.68 a | 23.36±3.89 b |
8/27 | 748.14±101.88 a | 760.15±84.70 a | 31.09±4.23 a | 22.39±3.07 b |
9/11 | 773.60±33.25 b | 807.09±103.92 a | 33.05±1.42 a | 25.01±3.50 b |
9/21 | 888.65±34.21 b | 919.22±70.34 a | 27.54±1.06 a | 25.53±2.94 b |
表10 库尔勒香梨及其芽变果实发育期果肉细胞长度及数量
Table 10 Pulp cell size and number during development of Korla pear and bud sport
采样日期 Sampling date(m/d) | 细胞长度 Cell size/μm | 细胞数量 Cell number/μm-2 | ||
---|---|---|---|---|
库尔勒香梨 Korla pear | 芽变 Bud sport | 库尔勒香梨 Korla pear | 芽变 Bud sport | |
6/13 | 183.22±32.53 b | 230.02±27.15 a | 20.07±3.56 a | 19.10±2.25 a |
6/28 | 257.08±30.09 b | 396.01±71.51 a | 25.22±2.95 a | 24.69±4.46 a |
7/13 | 426.63±35.74 b | 490.32±84.17 a | 31.07±2.60 a | 24.47±4.20 b |
7/28 | 512.87±67.87 b | 569.32±63.54 a | 21.37±2.83 b | 24.08±2.69 a |
8/12 | 579.04±81.59 b | 660.16±109.95 a | 26.15±3.68 a | 23.36±3.89 b |
8/27 | 748.14±101.88 a | 760.15±84.70 a | 31.09±4.23 a | 22.39±3.07 b |
9/11 | 773.60±33.25 b | 807.09±103.92 a | 33.05±1.42 a | 25.01±3.50 b |
9/21 | 888.65±34.21 b | 919.22±70.34 a | 27.54±1.06 a | 25.53±2.94 b |
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